(Citharinus_congicus ,(Citharinus_gibbosus ,((((((((Anostomus_ternetzi ,((Abramites_hypselonotus ,Leporinus_striatus ),(Laemolyta_proxima ,Schizodon_vittatus ))),((Caenotropus_labyrinthicus ,(Cyphocharax_spilotus ,Steindachnerina_insculpta )),(Prochilodus_lineatus ,Semaprochilodus_varii ))),((Apareiodon_affinis ,Parodon_nasus ),(Anodus_orinocensis ,(Hemiodus_thayeria ,Hemiodus_unimaculatus )))),(((Myloplus_rubripinnis ,((Metynnis_mola ,Acnodon_oligacanthus ),(Pristobrycon_striolatus ,(Serrasalmus_maculatus ,(Serrasalmus_gouldingi ,(Serrasalmus_rhombeus ,Pygocentrus_nattereri ,Serrasalmus_manueli )))))),(Mylossoma_duriventre ,(Piaractus_mesopotamicus ,Colossoma_macropomum ))),(Rhaphiodon_vulpinus ,(Cynodon_gibbus ,Hydrolycus_armatus )))),((Pyrrhulina_australis ,(Boulengerella_lateristriga ,Ctenolucius_hujeta )),(Hoplerythrinus_unitaeniatus ,(Hoplias_cf._malabaricus ,Hoplias_aimara )))),((((((Bryconetes_enigmaticus ,(((((((((((Acestrocephalus_sardina ,(Cynopotamus_argenteus ,(Galeocharax_humeralis ,Galeocharax_gulo ))),((Charax_leticiae ,Charax_stenopterus ),(Roeboides_descalvadensis ,Roeboides_microlepis ))),(Phenacogaster_calverti ,(Phenacogaster_franciscoensis ,Phenacogaster_tegatus ))),(Aphyocharacidium_bolivianum ,Axelrodia_lindeae ,Microschemobrycon_casiquiare ,Microschemobrycon_melanotus )),(Deuterodon_potaroensis ,(Moenkhausia_heikoi ,(Moenkhausia_doceana ,(Tetragonopterus_rarus ,((Tetragonopterus_argenteus ,(Tetragonopterus_carvalhoi ,(Tetragonopterus_chalceus ,Tetragonopterus_araguaiensis ))),(Tetragonopterus_anostomus ,Tetragonopterus_denticulatus ))))))),(Roeboexodon_guyanensis ,(Bryconexodon_juruenae ,Exodon_paradoxus ))),(((((((Acrobrycon_ipanquianus ,((Hemibrycon_dariensis ,(Hemibrycon_jabonero ,(Hemibrycon_plutarcoi ,(Hemibrycon_caucanus ,Hemibrycon_galvisi ,(Hemibrycon_boquiae ,Hemibrycon_brevispini ,Hemibrycon_quindos ))))),(((Hemibrycon_surinamensis ,(Hemibrycon_loisae ,(Hemibrycon_cristiani ,Hemibrycon_taeniurus ))),(Hemibrycon_helleri ,Hemibrycon_inambari )),(Hemibrycon_huambonicus ,(Hemibrycon_beni ,Hemibrycon_jelskii ))))),((Bryconamericus_pachacuti ,(((Ceratobranchia_obtusirostris ,((Attonitus_irisae ,(Attonitus_ephimeros ,Attonitus_bounites )),(Bryconacidnus_pectinatus ,Bryconamericus_diaphanus ))),(Rhinobrycon_negrensis ,((Knodus_gamma ,(Knodus_breviceps ,(Aulixidens_eugeniae ,Knodus_heteresthes ))),(((Knodus_alpha ,(Knodus_moenkhausii ,(Knodus_tanaothoros ,Knodus_orteguasae ))),(Knodus_meridae ,Knodus_deuterodonoides )),(Knodus_smithi ,(Knodus_megalops ,Knodus_cinarucoense )))))),((((Bryconamericus_microcephalus ,(Bryconamericus_patriciae ,(Odontostoechus_lethostigmus ,((Bryconamericus_agna ,Bryconamericus_uporas ),((Bryconamericus_iheringii ,(Bryconamericus_cf._iheringii ,Bryconamericus_rubropictus )),(Bryconamericus_ikaa ,Nantis_indefessus )))),(Hypobrycon_leptorhynchus ,Hypobrycon_maromba ))),((Diapoma_alburnus ,Diapoma_dicropotamicus ),(((Diapoma_tipiaia ,(Diapoma_obi ,(Hyphessobrycon_wajat ,Diapoma_guarani ))),(Diapoma_speculiferum ,Diapoma_terofali )),(Diapoma_uruguayense ,Diapoma_lepiclastus )))),(Piabarchus_analis ,(Bryconamericus_mennii ,(Bryconamericus_exodon ,Bryconamericus_stramineus )))),(Piabina_thomasi ,Piabina_argentea )))),((Microgenys_minuta ,(Carlastyanax_aurocaudatus ,(Creagrutus_anary ,(((Creagrutus_atrisignum ,(Creagrutus_taphorni ,((Creagrutus_muelleri ,(Creagrutus_melanzonus ,(Creagrutus_beni ,((Creagrutus_barrigai ,(Creagrutus_gephyrus ,Creagrutus_seductus )),(Creagrutus_britskii ,Creagrutus_ignotus ))))),(Creagrutus_peruanus ,(Creagrutus_ungulus ,(Creagrutus_occidaneus ,(Creagrutus_pila ,Creagrutus_changae ))))))),(Creagrutus_meridionalis ,(Creagrutus_affinis ,Creagrutus_brevipinnis ))),(Creagrutus_phasma ,(Creagrutus_cracentis ,Creagrutus_maxillaris )))))),(Lepidocharax_burnsi ,Planaltina_britskii )))),((Mimagoniates_microlepis ,(Mimagoniates_rheocharis ,Mimagoniates_inequalis )),(Glandulocauda_melanopleura ,Lophiobrycon_weitzmani ))),((Chrysobrycon_myersi ,(Pseudocorynopoma_doriae ,Pseudocorynopoma_heterandria )),(Corynopoma_riisei ,(Gephyrocharax_valencia ,(Gephyrocharax_atracaudatus ,Gephyrocharax_chocoensis ))))),(Argopleura_magdalenensis ,Argopleura_chocoensis )),(Tyttocharax_tambopatensis ,(Xenurobrycon_heterodon ,(Xenurobrycon_pteropus ,Xenurobrycon_polyancistrus )))),((Eretmobrycon_brevirostris ,((Eretmobrycon_emperador ,((Eretmobrycon_scleroparius ,Eretmobrycon_gonzalezi ),(Eretmobrycon_terrabensis ,Eretmobrycon_bayano ))),(Eretmobrycon_dahli ,(Eretmobrycon_peruanus ,Eretmobrycon_miraensis )))),(Hyphessobrycon_poecilioides ,Markiana_nigripinnis )))),((((Aphyocharax_nattereri ,((Aphyocharax_anisitsi ,(Aphyocharax_dentatus ,Aphyocharax_rathbuni )),(Aphyocharax_alburnus ,Aphyocharax_pusillus ))),(Prionobrama_filigera ,Prionobrama_paraguayensis )),(Paragoniates_alburnus ,(Phenagoniates_macrolepis ,(Xenagoniates_bondi ,Leptagoniates_steindachneri )))),(Protocheirodon_pi ,((((Heterocheirodon_yatai ,(Cheirodon_interruptus ,(Cheirodon_australe ,Cheirodon_kiliani ))),(Prodontocharax_melanotus ,(Saccoderma_melanostigma ,(Odontostilbe_dialeptura ,(Compsura_gorgonae ,Odontostilbe_mitoptera ))))),(((Odontostilbe_pequira ,(Odontostilbe_microcephala ,(Acinocheirodon_melanogramma ,Aphyocheirodon_hemigrammus ))),(((Odontostilbe_paraguayensis ,Odontostilbe_ecuadorensis ),(Odontostilbe_fugitiva ,(Compsura_heterura ,(Serrapinnus_microdon ,(Serrapinnus_calliurus ,Serrapinnus_notomelas ))))),(Odontostilbe_pulchra ,Odontostilbe_splendida ))),((Macropsobrycon_uruguayanae ,Cheirodon_stenodon ),(Kolpotocheirodon_figueiredoi ,Kolpotocheirodon_theloura )))),(Nanocheirodon_insignis ,(Pseudocheirodon_terrabae ,Pseudocheirodon_arnoldi )))))),((Hyphessobrycon_boulengeri ,(((((((((Aphyodite_grammica ,(Hemigrammus_unilineatus ,Moenkhausia_hemigrammoides )),((Hemigrammus_ulreyi ,((Ectrepopterus_uruguayensis ,Hyphessobrycon_heterorhabdus ),(Hemigrammus_bellottii ,Hemigrammus_schmardae ))),(Moenkhausia_collettii ,Moenkhausia_copei ))),(Hemigrammus_lunatus ,Hemigrammus_rodwayi )),(Moenkhausia_agnesae ,Moenkhausia_comma )),(Bryconella_pallidifrons ,(Pristella_maxillaris ,Pristella_ariporo ))),((Moenkhausia_inrai ,(Stichonodon_insignis ,(Brachychalcinus_copei ,(Stethaprion_erythrops ,(Orthospinus_franciscensis ,Poptella_paraguayensis ))))),(Gymnocorymbus_bondi ,(Gymnocorymbus_ternetzi ,Gymnocorymbus_thayeri )))),(((Nematocharax_venustus ,((Hasemania_nana ,(Hemigrammus_arua ,Moenkhausia_celibela )),(Moenkhausia_gracilima ,(Hemigrammus_erythrozonus ,(Hemigrammus_marginatus ,Moenkhausia_dichroura ))))),((Hyphessobrycon_meridionalis ,Hyphessobrycon_vanzolinii ),(Moenkhausia_lepidura ,((Moenkhausia_costae ,(Moenkhausia_bonita ,Moenkhausia_intermedia )),(Moenkhausia_jamesi ,Moenkhausia_lata ))))),(Moenkhausia_pirauba ,(Jupiaba_acanthogaster ,Moenkhausia_xinguensis )))),(((Brittanichthys_axelrodi ,(Paracheirodon_axelrodi ,Paracheirodon_innesi )),(Hemigrammus_bleheri ,Petitella_georgiae )),(Hyphessobrycon_compressus ,(Hyphessobrycon_pulchripinnis ,(Hyphessobrycon_eques ,(Hyphessobrycon_megalopterus ,(Hyphessobrycon_socolofi ,Hyphessobrycon_erythrostigma ))))))),(((((Astyanax_latens ,(((((Astyanax_moorii ,(Astyanax_lineatus ,(Astyanax_abramis ,Astyanax_lacustris ))),(Astyanax_chico ,Astyanax_troya )),(Astyanax_nasutus ,((Astyanax_bransfordii ,Astyanax_nicaraguensis ),(Astyanax_aeneus ,(Astyanax_baileyi ,Astyanax_caballeroi ,Astyanax_bailey ))))),(Hyphessobrycon_bifasciatus ,(Astyanax_paris ,(((Astyanax_cremnobates ,(Gymnocharacinus_bergii ,Hasemania_crenuchoides )),(Hyphessobrycon_anisitsi ,(Astyanax_eigenmanniorum ,(Astyanax_endy ,(Astyanax_puka ,Astyanax_rutilus ))))),(Oligosarcus_itau ,((Oligosarcus_pintoi ,(Oligosarcus_paranensis ,(Oligosarcus_bolivianus ,Oligosarcus_jenynsii ))),(Oligosarcus_longirostris ,Oligosarcus_menezesi ))))))),(Astyanax_correntinus ,(Astyanax_pelegrini ,(Ctenobrycon_hauxwellianus ,Psellogrammus_kennedyi ))))),(Hemigrammus_haraldi ,(Hemigrammus_pulcher ,Hemigrammus_aguaruna ))),((Hyphessobrycon_montagi ,(Hemigrammus_ataktos ,((Hemigrammus_coeruleus ,(Moenkhausia_cosmops ,(Bario_steindachneri ,(Moenkhausia_sanctaefilomenae ,(Moenkhausia_australe ,Moenkhausia_forestii ))))),(Moenkhausia_pyrophthalma ,(Thayeria_boehlkei ,Thayeria_obliqua ))))),((Hollandichthys_multifasciatus ,Rachoviscus_crassiceps ),(Hyphessobrycon_flammeus ,Hyphessobrycon_weitzmanorum )))),((Nematobrycon_palmeri ,(Pseudochalceus_kyburzi ,(Carlana_eigenmanni ,Rhoadsia_altipinna ))),(Grundulus_bogotensis ,Inpaichthys_kerri ))),((((Hemigrammus_levis ,Moenkhausia_ceros ),(Hyphessobrycon_herbertaxelrodi ,Erythrocharax_altipinnis )),((Moenkhausia_cotinho ,Macropsobrycon_xinguensis ),(Parapristella_georgiae ,Parecbasis_cyclolepis ))),(Hyphessobrycon_moniliger ,(Jupiaba_anteroides ,(Jupiaba_scologaster ,(Jupiaba_mucronata ,Jupiaba_polylepis )))))))),((Cyanogaster_noctivaga ,(Coptobrycon_bilineatus ,Hyphessobrycon_elachys )),(Stygichthys_typhlops ,(Hyphessobrycon_loweae ,(Phycocharax_rasbora ,(Myxiops_aphos ,(Probolodus_heterostomus ,(Deuterodon_sp. ,((Deuterodon_iguape ,(Deuterodon_langei ,Deuterodon_longirostris )),(Deuterodon_rosae ,Deuterodon_stigmaturus ))))))))))),(Atopomesus_pachyodus ,(Amazonspinther_dalmata ,(Spintherobolus_ankoseion ,Spintherobolus_papilliferus )))),(Paleotetra_entrecorregos ,Paleotetra_aiuruoca ))),((Lignobrycon_myersi ,((Clupeacharax_anchoveoides ,(Agoniates_anchovia ,Agoniates_halecinus )),(Triportheus_nematurus ,(Triportheus_angulatus ,Triportheus_pantanensis )))),(Engraulisoma_taeniatum ,(Thoracocharax_stellatus ,(Carnegiella_strigata ,Carnegiella_marthae ))))),(Brycon_henni ,((Brycon_pesu ,((Brycon_falcatus ,Brycon_polylepis ,Henochilus_wheatlandii ),(Brycon_moorei ,(Brycon_gouldingi ,Brycon_orbignyanus )))),(Salminus_franciscanus ,(Salminus_brasiliensis ,Salminus_hilarii ))))),(((Acestrorhynchus_microlepis ,(Acestrorhynchus_falcatus ,(Acestrorhynchus_lacustris ,Acestrorhynchus_pantaneiro ))),((Gnathocharax_steindachneri ,(Hoplocharax_goethei ,(Heterocharax_macrolepis ,Lonchogenys_ilisha ))),(Gilbertolus_maracaiboensis ,Roestes_ogilviei ))),((Bryconops_caudomaculatus ,(Bryconops_affinis ,Bryconops_melanurus )),(Iguanodectes_geisleri ,Piabucus_melanostoma )))),(Chalceus_macrolepidotus ,Chalceus_epakros )),(Arnoldichthys_spilopterus ,(((Hydrocynus_goliath ,(Alestes_macrophthalmus ,(Brachyalestes_carolinae ,(Bryconalestes_longipinnis ,(Bryconaethiops_macrops ,Alestopetersius_hilgendorfi ))))),(Micralestes_stormsi ,Micralestes_acutidens )),((Hepsetus_odoe ,Hepsetus_kingsleyae ),(Hepsetus_occidentalis ,(Hepsetus_lineatus ,(Hepsetus_cuvieri ,Hepsetus_microlepis )))))))),(Crenuchus_spilurus ,(Characidium_zebra_gr. ,Characidium_rachovii ))),(Xenocharax_spilurus ,(Nannocharax_uniocellatus ,((Distichodus_fasciolatus ,(Distichodus_maculatus ,Distichodus_decemmaculatus )),(Eugnathichthys_macroterolepis ,(Mesoborus_crocodilus ,(Belonophago_hutsebouti ,Phago_boulengeri )))))))));